The ontological primacy of umwelt

Where do the basic composite parts of an umwelt – organism and environment come from? Customarily, in theory, emphasis is on their mutual co-construction or constitution through functional cycles. But another question could be added to this inquiry: what origin, genesis organisms environments which are now indivisibly united once again? On basis transactional conception relations proposed by John Dewey, concept schismogenesis as described Gregory Bateson, process individuation explicated Gilbert Simondon, it that ontologically primary, its internal division, separation individuation, results appearance with environments.